Dr. Werner Heitland

Chair for Terrestrial Ecology

Department of Ecology and Ecosystemmanagement
Technische Universität München
Hans-Carl-von-Carlowitz-Platz 2
D-85350 Freising-Weihenstephan

Phone:  +49.8161.71.4596
Fax:      +49.8161.71.4427
E-mail: heitland[at]forst.tu-muenchen.de

Curriculum vitae

1991- 2011 Assistant at the Department of Animal Ecology, TUM.
1991 PhD at the Department of Ecology (University of Kiel) about Biology, Ecology and Parasitoids of Platycampus luridiventris. (Hym., Tenthr.) under Prof. Dr. Hubert Pschorn-Walcher.
Study of biology and chemistry at the University of Bochum and Kiel.

Research interest

  • Horse chestnut leaf-miner (Cameraria ohridella)
  • Interaction between plants, phytophagous insects and their parasitoids.
  • Sawflies and their parasitoid complexes.
  • Feeding-strategies of phytophagous insects.



Kopelke, J.-P., W. Heitland, and E. Altenhofer. 2012. Survival and Mortality Factors in two Central European Populations of Euura amerinae Stem gallers (Hymenoptera: Tenthredinidae: Nematinae). Entomologia Generalis 34:33-60. PDF

Utschick, H., S. C. Michl, and W. Heitland. 2012. Indikationspotenzial von Auwaldvögeln für das Monitoring von Auenrenaturierungen. Ornithol. Anz. 51:97-134. PDF


Augustin, S., S. Guichard, W. Heitland, J. Freise, A. Svatoš, and M. Gilbert. 2009. Monitoring and dispersal of the invading Gracillariidae Cameraria ohridella. Journal of Applied Entomology 133:58-66.


Menzel, A., N. Estrella, W. Heitland, A. Susnik, C. Schleip, and V. Dose. 2008. Bayesian analysis of the species-specific lengthening of the growing season in two European countries and the influence of an insect pest. International Journal of Biometeorology 52:209-218.


Hedenström, E., H. Edlund, A.-B. Wassgren, G. Bergström, O. Anderbrant, F. Östrand, A. Sierpinski, M.-A. Auger-Rozenberg, A. Herz, W. Heitland, and M. Varama. 2006. Sex Pheromone of the Pine Sawfly, Gilpinia pallida : Chemical Identification, Synthesis, and Biological Activity. Journal of Chemical Ecology 32:2525-2541.


Anderbrant, O., F. Östrand, G. Bergström, A.-B. Wassgren, M.-A. Auger-Rozenberg, C. Geri, E. Hedenström, H.-E. Högberg, A. Herz, and W. Heitland. 2005. Release of sex pheromone and its precursors in the pine sawfly Diprion pini (Hym., Diprionidae). Chemoecology 15:147-151.

Gilbert, M., S. Guichard, J. Freise, J. C. Gregoire, W. Heitland, N. Straw, C. Tilbury, and S. Augustin. 2005. Forecasting Cameraria ohridella invasion dynamics in recently invaded countries: from validation to prediction. Journal of Applied Ecology 42:805-813.

Heitland, W. and H. Pscorn-Walcher. 2005. Biology and parasitoids of the peculiar alder sawfly, Platycampus luridiventris (Fallen) (Insecta, Hymenoptera, Tenthredinidae). Senckenbergiana Biologica 85:215-231.

Herz, A. and W. Heitland. 2005. Species diversity and niche separation of cocoon parasitoids in different forest types with endemic populations of their host, the common pine sawfly Diprion pini (Hymenoptera : Diprionidae). European Journal of Entomology 102:217-224.

Werner Heitland and J. Freise. 2005. Die Rosskastanien-Miniermotte, der auffälligste Schädling der weißblühenden Rosskastanie. LWF Wissen 48:28-30.


Freise, J. F. and W. Heitland. 2004. Bionomics of the horse-chestnut leaf miner Cameraria ohridella Deschka & Dimic 1986, a pest on Aesculus hippocastanum in Europe (Insecta, Lepidoptera, Gracillariidae). Senckenbergiana Biologica 84:61-80.

Gilbert, M., J. C. Grégoire, J. F. Freise, and W. Heitland. 2004. Long-distance dispersal and human population density allow the prediction of invasive patterns in the horse chestnut leafminer Cameraria ohridella. Journal of Animal Ecology 73:459-468.


Baumann, R., R. Schubert, W. Heitland, M. A. Auger-Rozenberg, P. Faivre-Rimpant, and G. Müller-Starck. 2003. Genetic diversity within and among populations of Diprion pini (Hym., Diprionidae) determined by random amplified polymorphic DNA-polymerase chain reaction of haploid males. Journal of Applied Entomology 127:258-264.

Freise., J. and W. Heitland. 2003. The Waipuna Hot Foam System: a chance for Cameraria ohridella control? Nachrichtenblatt des Deutschen Pflanzenschutzdienstes 55:242-244.

Herz, A. and W. Heitland. 2003. Impact of cocoon predation and parasitism on endemic populations of the common pine sawfly, Diprion pini (L.) (Hymenoptera, Diprionidae) in different forest types. Agricultural and Forest Entomology 5:35-42.

Thalmann, C., J. Freise, W. Heitland, and S. Bacher. 2003. Effects of defoliation by horse chestnut leafminer (Cameraria ohridella) on reproduction in Aesculus hippocastanum. Trees - Structure and Function 17:383-388.


Freise, J. F., W. Heitland, and I. Tosevski. 2002. Parasitism of the horse chestnut leaf miner, Cameraria ohridella Deschka and Dimic (Lep., Gracillariidae), in Serbia and Macedonia. Anzeiger Fur Schadlingskunde-Journal of Pest Science 75:152-157.

Jiri Kindl, Blanka Kalinova, Jona Freise, Werner Heitland, Sylvie Augustin, Sylvain Guichard, Nikos Avtzis, and A. Svatos. 2002. Monitoring the Population Dynamics of the Horse Chestnut Leafminer Cameraria ohridella with a Synthetic Pheromone in Europe. Plant Protection Science 38:131-138.

1988 bis 2000

Herz, A., W. Heitland, O. Anderbrant, H. Edlund, and E. Hedenström. 2000. First use of pheromones to detect phenology patterns and density relationships of pine sawflies in German forests. Agricultural and Forest Entomology 2:123-129.

Freise, J. and W. Heitland. 1999. A brief note on sexual differences in pupae of the horse-chestnut leaf miner, Cameraria ohridella Deschka & Dimic (1986) (Lep., Gracillariidae), a new pest in Central Europe on Aesculus hippocastanum. Journal of Applied Entomology 123:191-192.

Herz, A. and W. Heitland. 1999. Larval parasitism of a forest pest, the common pine sawfly Diprion pini (L.) (Hym., Diprionidae), during an endemic density phase. Journal of Applied Entomology 123:129-137.

Schroeter, B., M. Olech, L. Kappen, and W. Heitland. 1995. Ecophysiological investigations of Usnea anfarcficain the maritime Antarctic 1. Annual microclimatic conditions and potential primary production. Antarctic Science 7:251-260.

Herbst, J. and W. Heitland. 1994. Genetic differentiation among populations of the sawfly-species Platycampus luridiventris, associated with different alder species (Hymenoptera: Tenthredinidae). Entomologia Generalis 19:39-48.

Heitland, W. and H. Pschorn-Walcher. 1992. A brief note on unusually high superparasitism in some sawfly parasitoids (Hym., Ichneumonidae: Euryproctini). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 65:75-79.

Heitland, W. and H. Pschornwalcher. 1992. Biological differences between populations of Platycampus luridiventris feeding on different species of alder (Hymenoptera : Tenthredinidae). Entomologia Generalis 17:185-194.

Heitland, W. 1988. Untersuchungen an Parasitoiden von Dipteren im Strandanwurf der Kieler Förde. Bonn. zool. Beitr. 39:129-145.